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Wolf sable/gray, Wolf sable, Wolf gray

The A Locus (Agouti series) interacts closely with the E, K, and B loci which can lead to a dog’s overall coat color and pattern. A-Locus mutations are only expressed if the dog is “Clear” or “Carrier” at the E locus and “Clear” at the K-KB locus. Four potential mutations at the A Locus can each have a different effect on coat color. The mutations are known as A-AY, A-AW, A-AT, and A-A and can determine whether a dog is a Carrier of sable/fawn, wolf sable/gray, black and tan/tricolor/tan points coloration or a recessive form of a solid black or bicolor coat color. The A-AW Locus mutation results in a dog that is wolf-sable/gray.

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These dogs have two copies of the normal gene and the effect of A Locus on their coat color can be determined by testing the A-AY, A-AT, and A-A loci. They will also not pass this mutation to any of their offspring.


These dogs have one copy of the normal gene and one copy of the mutation associated with this coat color. They will develop a wolf sable/gray coat due to this mutation in the absence of the AY mutation. They will, if bred, pass the mutation to 50% of their offspring, on average.


These dogs have two copies of the mutation and will typically develop a wolf gray/sable coat due to the A-AW Locus mutation and will pass this mutation to 100% of their offspring. However, this dog’s coat color is also dependent on the E, K, and B Locus mutations.

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Additional Details


Complex order of dominance with other A Locus mutations (AY > AW > AT > A) and also depends on results for E, K, and B Locus mutations.

Affected gene



Ch. 24


Clear for SINE insertion of approximately 239 base pairs in intron 1a and c.244G>T and 248G>A


Kerns JA, Newton J, Berryere TG, Rubin EM, Cheng JF, Schmutz SM, Barsh GS. Characterization of the dog Agouti gene and a nonagouti mutation in German Shepherd Dogs. Mamm Genome. 2004 Oct; 15(10):798-808. [PubMed: 15520882]